Luke Jacobus, Ph.D.

I grew up in the Columbus, IN, area and have returned to this community. Currently, my students and I investigate various topics in environmental science, especially those related to freshwater insects. These organisms have proven valuable for the development of new generation identification systems, the analysis of biodiversity patterns in space & time, and the application of phylogenetic models to problems in environmental toxicology. Much of my research has focused on the insect order Ephemeroptera, or mayflies. Aspects of this research include, but are not limited to, taxonomy and identification, DNA barcoding, biodiversity, phylogeny, rearing and stage associations, descriptive biology and ecology, field studies, conservation, species inventories and regional faunistics. I interact with various international stakeholders in the academic, government and private realms.

• Bachelor of Science, with highest distinction, Entomology. Minor, International Studies in Agriculture. Purdue University, West Lafayette, Indiana. May 2000.
• Doctor of Philosophy, Entomology. Dissertation title: Phylogeny and Classification of the Mayfly Subfamily Ephemerellinae (Ephemeroptera: Ephemerellidae). Purdue University, West Lafayette, Indiana; W.P. McCafferty, advisor. August 2006.

• Biological Survey of Canada
• Indiana Academy of Science
• Association of College & University Biology Educators (ACUBE)
• Society for Freshwater Science
• Entomological Society of America
• Phi Beta Kappa
• Carolina Area Benthologists Association

• BIOL-K 101 Concepts of Biology I
• BIOL-K 103 Concepts of Biology II
• BIOL-K 295 Special Assignments
• BIOL-K 341 Principles of Ecology & Evolution
• BIOL-K 342 Ecology Laboratory
• BIOL-K 395 Advanced Topics in Biology
• BIOL-K 411 Global Change Biology
• BIOL-K 490 Capstone in Biology
• BIOL-K 493 Independent Research
• BIOL-K 494 Senior Research Thesis
• BIOL-N 100 Contemporary Biology
• BIOL-N 108 Plants, Animals & the Environment

• Faculty Senate President
• Interim Director, IUPUC Honors Program
• Campus Sustainability Committee, IUPUC
• Senior Editor, BMC Ecology amd Evolution
• Subject Editor, Zootaxa
• Editorial Board, Bioscene: Journal of College Biology Teaching
• Visiting Scholar, Department of Entomology, Purdue University
• Research Associate, Purdue University Entomological Research Collection
• Steward, Mayfly Central

Mayflies are thought to be the oldest extant group of winged insects. What stories they must have to tell! Little by little, we are piecing together their part of the Earth's natural history. To get the full picture, we will need to answer several questions. Which species are here? What do we call them? Where do they live? How do they function? How do they interact? Gathering information about species' geographic distributions, biologies and morphologies (from external body structure to DNA sequence) will help answer these basic questions. Of further importance, mayfly species are commonly utilized worldwide as indicators of environmental health and are often the subjects of ecological and physiological studies.

• 2022: Indianapolis Zoo Short-term Conservation Grant
• 2016-2021: Collections in Support of Biological Research (Natural History) Grant, National Science Foundation
• 2021, 2016, 2014: Outstanding Fulltime Faculty Research, Scholarship and Creativity Award, IUPUC
• 2022: Hoosier RiverWatch Instructor
• 2019-present: Muscatatuck National Wildlife Refuge volunteer
• 2019-2021, 2016-2017: Consultant, Indiana Natural Heritage Data Center Indiana Department of Natural Resources Nature Preserves
• 2019-2021: Duke Energy Foundation Grant.
• 2022, 2015: Consultant, General Studies of Species in Canada, Canadian Wildlife Service
• 2017: Hoosier RiverWatch workshop
• 2015, 2017: Southeast Water Pollution Biologists Association Workshop
• 2014: Fellow, Indiana Academy of Science
• 2013: Southwestern Association of Freshwater Invertebrate Taxonomists workshop, California State University, Long Beach
• 2013, 2015: Carolina Area Benthologists Workshop, Hot Springs, North Carolina
• 2013-present: Campus Constitution & Citizenship Day co-coordinator
• 2012: Senior Research Grant, Indiana Academy of Science
• 2005: Northwest Biological Assessment Workgroup workshop, University of Idaho, Moscow
• 2003-06: National Science Foundation Graduate Research Fellow
• 2001-03: USEPA STAR Graduate Research Fellow
• 2000: Outstanding Senior in Agriculture & Veterinary Medicine, Purdue University
• 2000. Student Response Commencement Address, Purdue University

• Harman, J., C. A. Hipsley, L. M. Jacobus, D. A. Liberles, J. Settele, and A. Traulsen. 2022. 2022 BMC Ecology and Evolution image competition: the winning images. BMC Ecology and Evolution 22: 99. https://doi.org/10.1186/s12862-022-02049-y
• Jacobus, L. M., M. Sartori, C. Molineri, T. Tiunova, and L. Pereira. 2022. Zootaxa Ephemeroptera editors' annual summary and acknowledgments (2021). The Mayfly Newsletter 25(1): 11.
• Li, X.-F., Y.-K. Sun, Z.-Y. Liu, L. M. Jacobus, and W. Xiao. 2022. A new species of Notacanthella Jacobus & McCafferty, 2008 (Ephemeroptera: Ephemerellidae) from Yunnan, China. ZooKeys 1103: 25-44. https://doi.org/10.3897/zookeys.1103.82984
• Tungpairojwong, N., S. Phlai-Ngam, and L. M. Jacobus. 2022. A new species of Acentrella Bengtsson, 1912 (Ephemeroptera: Baetidae) from Thailand. Zootaxa 5125(4): 351-378. https://doi.org/10.11646/zootaxa.5125.4.1
• Jacobus, L. M. 2022. Reassessment of the conservation status of Ephemeroptera in YT, NT, NU, BC, AB, SK, MB, ON, QC, NB, NS, PE, LB, and NF. General Status of Species in Canada, Canadian Wildlife Service, Environment and Climate Change Canada, Gaineau, Quebec. 3 databases.
• Srinivasan, P., T. Sivaruban, S. Barathy, R. Isack, and L. M. Jacobus. 2022. A new species of Clypeocaenis Soldan 1978 (Ephemeroptera: Caenidae) from Tamil Nadu, India. Zootaxa 5091(3): 467-476. https://doi.org/10.11646/zootaxa.5091.3.6
• Jacobus, L. M. 2021. Editorial: Current contributions to the systematics of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae). Zootaxa 5076: 4-6.
• Jacobus, L. M. (Ed.) 2021. Current contributions to the systematics of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae). Zootaxa 5076. Magnolia Press, Auckland, New Zealand. 70 pp.
• Jacobus, L. M. 2021. Profiles: Lillien Jane Martin (1851-1943). Indiana Academy of Science Newsletter 181: 7.
• Jacobus, L. M. 2021. A note from the editor. Indiana Academy of Science Newsletter 181: 12.
• Harman, J. L., A. L. Cuff, J. Settele, L. M. Jacobus, D. A. Liberles, and A. Traulsen. 2021. Correction to: Inaugural BMC Ecology and Evolution image competition: the winning images. BMC Ecology and Evolution 21: 174.
• Harman, J. L., A. L. Cuff, J. Settele, L. M. Jacobus, D. A. Liberles, and A. Traulsen. 2021. Inaugural BMC Ecology and Evolution image competition: the winning images. BMC Ecology and Evolution 21: 157.
• Jacobus, L. M., M. Sartori, F. F. Salles, L. Pereira, C. Molineri, and T. Tiunova. 2021. Zootaxa Ephemeroptera Editors' Annual Summary and Acknowledgements (2020). The Mayfly Newsletter 24(1): 8.
• Jacobus, L. M., F. F. Salles, B. Price, L. Pereira-da-Conceicoa, E. Dominguez, P. J. Suter, C. Molineri, T. Tiunova, and M. Sartori. 2021. Mayfly taxonomy (Arthropoda: Hexapoda: Ephemeroptera) during the first two decades of the twenty-first century and the concentration of taxonomic publishing. Zootaxa 4979(1): 25-30. https://doi.org/10.11646/zootaxa.4979.1.6
• Martynov, A. V., C. Selvakumar, D. M. Palatov, K. A. Subramanian, K. G. Sivaramakrishnan, M. Vasanth, and L. M. Jacobus. Overview of Indian and Nepali representatives of the Cincticostella nigra (Ueno, 1928) complex (Ephemeroptera: Ephemerellidae), with discussion about Cincticostella Allen, 1971 species complexes. ZooKeys 1040: 123-166. doi: 10.3897/zookeys.1040.64280
• Martynov, A. V., C. Selvakumar, K. A. Subramanian, K. G. Sivaramakrishnan, M. Vasanth, B. Sinha, and L. M. Jacobus. 2021. Overview of Indian Hyrtanellini (Ephemeroptera: Ephemerellidae), with new species and records from related regions. Zootaxa 4975(3): 451-482. https://doi.org/10.11646/zootaxa.4975.3.2
• Li, X.-F., S.-C. Ao, X. Shi, J.-J. Cheng, L. Tan, Q.-H. Cai, and L. M. Jacobus. 2021. Life history of Caenis lubrica Tong and Dudgeon, 2002 (Ephemeroptera: Caenidae) in a Three Gorges Reservoir feeder stream, subtropical Central China. Aquatic Insects 42(1): 50-61, DOI: 10.1080/01650424.2020.1867869.
• Jacobus, L. M. 2021. The type locales of Heptagenia whitingi Webb & McCafferty, 2007 and Kirmaushenkreena zarankoae McCafferty, 2011 (Insecta: Ephemeroptera: Heptageniidae, Baetidae). Proceedings of the Indiana Academy of Science (2020) 129(1): 65.
• Jacobus, L. M. 2021. Indiana's big river mayflies (Insecta: Ephemeroptera): Final report on 2019-2020 sampling. Report, Indiana Natural Heritage Data Center, Division of Nature Preserves, Indiana Department of Natural Resources, Indianapolis, Indiana. 20 pp. + spreadsheet.
• Jacobus, L. M. 2021. Erratum: Nikita J. Kluge (2020) Demoulinia Gillies 1990 and two unnamed genera of the plesiomorphon Protopatellata (Ephemeroptera, Baetidae) from South Africa. Zootaxa, 4820, 438-464. Zootaxa 4903(4): 600.
• Jacobus, L. M. 2020. Zootaxa update. The Mayfly Newsletter 23(2): 5.
• Auychinda, C., L. M. Jacobus, M. Sartori, B. Boonsoong. A new species of Vietnamella Tshernova 1972 (Ephemeroptera: Vietnamellidae) from Thailand. Insects 11(9):554. https://doi.org/10.3390/insects11090554
• Cuff, A. L., M. Baguette, S. Blanchet, L. M. Jacobus, D. Mazzi, and J. Settele. 2020. Seventh BMC ecology image competition: the winning images. BMC Ecology 20 (42). https://doi.org/10.1186/s12898-020-00310-w.
• Jacobus, L. M. 2020. Indiana's big river mayflies (Insecta: Ephemeroptera): July & August 2019 sampling. Proceedings of the Indiana Academy of Science [2019] 128(2): 267.
• Jacobus, L. M. 2020. Some notable records of mayflies (Insecta: Ephemeroptera) from big rivers in Indiana. BMC Research Notes 13 (357). https://doi.org/10.1186/s13104-020-05198-9.
• Jacobus, L. M. 2020. Some Notable Records of Mayflies (Insecta: Ephemeroptera) from Big Rivers in Indiana: Supporting Data (Version 08.vi.2020) [Data set]. Zenodo. http://doi.org/10.5281/zenodo.3885370.
• Jacobus, L. M. 2020. Some Notable Records of Mayflies (Insecta: Ephemeroptera) from Big Rivers in Indiana: Methodology (Version 08.vi.2020) [Data set]. Zenodo. http://doi.org/10.5281/zenodo.3885417.
• Zaspel, J. M., J. M. Allen, C. D. Tyrrell, N. Lemoine, L. M. Jacobus, C. Klem, J. Goodwin, and J. Bates. 2020. Human health, interagency coordination, and the need for biodiversity data. BioScience 70(7): 527.
• Jacobus, L. M., M. Sartori, F. F. Salles, and P. Suter. 2020. Zootaxa Ephemeroptera editors' annual summary and acknowledgments (2019). The Mayfly Newsletter 23(1): 7.
• Jacobus, L. M. 2020. Biodiversity research and the impacts of a global pandemic response. BMC Series Blog, 16 June 2020. https://blogs.biomedcentral.com/bmcseriesblog/2020/06/16/biodiversity-research-and-the-impacts-of-a-global-pandemic-response [Reprinted from: BMC Series Editorial Site, Covid-19: Featured blogs and resources, 27 May 2020.]
• Jacobus, L. M. 2020. Biodiversity research and the impacts of a global pandemic response. BMC Series Editorial Site, Covid-19: Featured blogs and resources, 27 May 2020. https://bmcserieseditors.biomedcentral.com/users/117002-luke-m-jacobus/posts/biodiversity-research-and-the-impacts-of-a-global-pandemic-response.
• Jacobus, L. M. 2020. Profiles: William Albert Noyes (1857-1941). Indiana Academy of Science Newsletter 175: 7-8.
• Jacobus, L. M. 2019. Profiles: John Casper Branner 1850-1922. Indiana Academy of Science Newsletter 173: 7.
• Jacobus, L. M. 2019. Indiana's big river mayflies (Insecta: Ephemeroptera), Part 1: July & August 2019 sampling. Report, Indiana Natural Heritage Data Center, Division of Nature Preserves, Indiana Department of Natural Resources, Indianapolis, Indiana. 15 pp. + spreadsheet.
• Bolton, M. J., S. K. Macy, R. E. Dewalt, and L. M. Jacobus. 2019. New Ohio and Indiana records of aquatic insects (Ephemeroptera, Plecoptera, Trichoptera, Coleoptera: Elmidae, Diptera: Chironomidae). Ohio Biological Survey Notes 9: 1-15.
• Green, M. W., P. W. Blum, S. C. Sellers, M. M. Gangloff, L. M. Jacobus and S. R. Tuberty. 2019. Mesohabitat current velocity effects on Didymosphenia geminata and macroinvertebrates in a SE USA hypolimnetic tailwater. Aquatic Ecology 53(4): 607-628. https://doi.org/10.1007/s10452-019-09712-5.
• Jacobus, L. M. 2019. Remembering Marion T. Jackson (19 Aug 1933-7 Feb 2019). Indiana Academy of Science Newsletter 172: 5.
• Jacobus, L. M., M. Sartori, F. F. Salles, and P. Suter. 2019. Zootaxa Ephemeroptera editors' annual summary and acknowledgments (2018). The Mayfly Newsletter 22(1):3.
• Jacobus, L. M., C. R. Macadam, and M. Sartori. 2019. Mayflies (Ephemeroptera) and their contributions to ecosystem services. In: Morse JC, Adler PH (Eds) Diversity and Ecosystem Services of Aquatic Insects. Insects 10(6), 170: 1-26. https://doi.org/10.3390/insects10060170
• Martynov, A. V., C. Selvakumar, K. A. Subramanian, K. G. Sivaramakrishnan, K. Chandra, D. M. Palatov, B. Sinha, and L. M. Jacobus. 2019. Review of the Cincticostella insolta (Allen, 1971) complex (Ephemeroptera: Ephemerellidae), with description of three new species from northern India and Nepal. Zootaxa 4551(2): 147-179. https://doi.org/10.11646/zootaxa.4551.2.2
• Jacobus, L. M. 2019. Ephemeroptera of Canada. In: Langor DW, Sheffield CS (Eds) The Biota of Canada - A Biodiversity Assessment. Part 1: The Terrestrial Arthropods. ZooKeys 819: 211-225. doi: 10.3897/zookeys.819.26411
• Cuff, A. L., Y. Lou, J. Zhigang, M. Baguette, S. Blanchet, J. Clobert, L. M. Jacobus, D. Mazzi and J. Settele. 2019. BMC ecology image competition 2018: the winning images. BMC Ecology 19:11.
• Jacobus, L. M. 2019. Feature photo: Hexagenia sp. (Ephemeridae) in the morning dew. The Mayfly Newsletter 21(2): 1.
• Jacobus, L. M. 2018. Endemism among Canada's mayfly species (Insecta: Ephemeroptera). Newsletter of the Biological Survey of Canada 37(1): 13-19.
• Jacobus, L. M. 2018. Endemism among Canada's mayfly species (Insecta: Ephemeroptera). The Mayfly Newsletter 21(1): 13-19. [Reprinted with permission from Newsletter of the Biological Survey of Canada]
• Jacobus, L. M. 2018. Mayfly Central expands coverage. The Mayfly Newsletter 21(1): 20.
• Jacobus, L. M., M. Sartori, F. F. Salles and P. Suter. 2018. Zootaxa Ephemeroptera editors' annual summary and acknowledgments (2017), with special remarks about preparing manuscripts. The Mayfly Newsletter 21(1): 10.
• Webb, J. M., L. M. Jacobus, and S. P. Sullivan. 2018. The state of systematics of North American Baetis Leach, 1815 (Ephemeroptera: Baetidae), with recommendations for identification of larvae. Zootaxa 4394(1): 105-127.
• Jacobus, L. M. 2017. Identifying larvae of eastern North American mayfly species (Insecta: Ephemeroptera). Southeast Water Pollution Biologists Association, Grand Rivers, Kentucky. 29 pp. Unpublished workshop manual.
• Jacobus, L. M. 2017. Conservation status rankings of Indiana mayflies (Insecta: Ephemeroptera). Report, Indiana Natural Heritage Data Center, Division of Nature Preserves, Indiana Department of Natural Resources, Indianapolis, Indiana. Spreadsheet.
• Jacobus, L. M., and P. Harper. 2017. Mayflies: Common names. Report, General Status of Species in Canada, Canadian Wildlife Service, Environment & Climate Change Canada, Government of Canada. Spreadsheet.
• Foote, C., C.T. Darimont, M. Baguette, S. Blanchet, L.M. Jacobus, D. Mazzi, and J. Settele. 2017. BMC Ecology Image Competition 2017: the winning images. BMC Ecology 17: 28. Available online: https://bmcecol.biomedcentral.com/articles/10.1186/s12898-017-0138-8.
• Rogers, D.C., S.T. Ahyong, C.B. Boyko, C.D. D'Acoz, et al. 2017. Images are not and should not ever be type specimens: a rebuttal to Garraffoni & Freitas. Zootaxa 4269(4): 455-459.
• Cruz, P.V., A.K. Usher and L.M. Jacobus. 2017. Review of continental North and Central American Paracloeodes Day 1955 (Ephemeroptera: Baetidae), including description of a new species from Minnesota, U.S.A. Zootaxa 4263(2): 333-346.
• Morse, J.C., B.P. Stark, and L.M. Jacobus. 2017. Chapter 1: Introduction. Pp. 10-14, in: J.C. Morse, W.P. McCafferty, B.P. Stark, and L.M. Jacobus (Eds). Larvae of the Southeastern USA Mayfly, Stonefly, and Caddisfly Species (Ephemeroptera, Plecoptera, and Trichoptera). Biota of South Carolina, Vol. 9. Clemson University Public Service Publishing, Clemson University, Clemson, South Carolina, USA. 482 pp.
• McCafferty, W.P., L.M. Jacobus, A.V. Provonsha, and N.A. Wiersema. 2017. Chapter 2: Ephemeroptera. Pp. 15-160, in: J.C. Morse, W.P. McCafferty, B.P. Stark, and L.M. Jacobus (Eds). Larvae of the Southeastern USA Mayfly, Stonefly, and Caddisfly Species (Ephemeroptera, Plecoptera, and Trichoptera). Biota of South Carolina, Vol. 9. Clemson University Public Service Publishing, Clemson University, Clemson, South Carolina, USA. 482 pp.
• Morse, J.C., W.P. McCafferty, B.P. Stark, and L.M. Jacobus (Eds). 2017. Larvae of the Southeastern USA Mayfly, Stonefly, and Caddisfly Species (Ephemeroptera, Plecoptera, and Trichoptera). Biota of South Carolina, Vol. 9. Clemson University Public Service Publishing, Clemson University, Clemson, South Carolina, USA. 482 pp. https://secure.touchnet.net/C20569_ustores/web/store_cat.jsp?STOREID=168&CATID=352&SINGLESTORE=true
• Bright, E. and L. M. Jacobus. 2017. Book Review: Mayfly Larvae of Wisconsin, by Tom H. Klubertanz (2016). The Mayfly Newsletter 20(1): 8-9.
• Jacobus, L. M., M. Sartori, F. F. Salles, and P. Suter. 2017. Zootaxa Ephemeroptera Editors' Annual Summary and Acknowledgments (2016). The Mayfly Newsletter 20(1): 3.
• Jacobus, L. M., M. Sartori, and F. F. Salles. 2016. Zootaxa Ephemeroptera editors' annual summary and acknowledgments (2015). The Mayfly Newsletter 19(2): 3.
• Jacobus, L. M., M. Sartori, F. Salles, and P. Suter. 2016. New editorial structure for Zootaxa Ephemeroptera submissions. The Mayfly Newsletter 19(2): 3
• Jacobus, L. M. 2016. News from Mayfly Central. The Mayfly Newsletter 19(2): 4.
• Jacobus, L. M. 2016. Importance of entomological collections. The Mayfly Newsletter 19(2): 4.
• Jacobus, L. M. 2016. An annotated list of Indiana mayflies (Insecta: Ephemeroptera). Report, Indiana Natural Heritage Data Center, Division of Nature Preserves, Indiana Department of Natural Resources, Indianapolis, Indiana. 21 pp.
• Salles, F. F., K. B. Angeli, and L. M. Jacobus. 2016. Review of Waltzoyphius McCafferty & Lugo-Ortiz, 1995 and Zelusia Lugo-Ortiz & McCafferty, 1998 (Ephemeroptera: Baetidae). Zoosymposia 11: 174-204.
• Simundza, J., M. Palmer, J. Settele, L.M. Jacobus, D.P. Hughes, D. Mazzi and S. Blanchet. 2016. BMC Ecology Image Competition 2016: the winning images. BMC Ecology 16(34). Available online: https://bmcecol.biomedcentral.com/articles/10.1186/s12898-016-0090-z
• Jacobus, L. M. 2016. Lepidophora lutea Painter, 1962 (Insecta: Diptera: Bombyliidae: Ecliminae). Proceedings of the Indiana Academy of Science 124(2).
• Holland, V. B., S. R. Beaty, and L. M. Jacobus. 2016. A new species of Neoephemera McDunnough, 1925 (Ephemeroptera: Neoephemeridae) from North Carolina and Virginia. Zootaxa 4138: 139-154.
• Parker, C. H. & Jacobus, L. M. 2016. Richard Henry Daggy, Ph.D., Dr.P.H. (1914-2001). The Mayfly Newsletter 19(1): 6-7.
• Jacobus, L. M. 2016. Purdue Type Collections. The Mayfly Newsletter 19(1): 7.
• Jacobus, L. M. 2016. Addendum to Book Review of The Mayflies of Europe, by Bauernbeind & Soldan (2012), Published in American Entomologist 61(3), DOI: 10.1093/ae/tmv056. The Mayfly Newsletter 19(1): 8-9.
• Jacobus, L. M. 2016. 2014 Ephemeroptera Bibliography. The Mayfly Newsletter 19(1): 10-12.Jacobus, L. M. 2016. Pp. 51-59, in: Wetzel, M. J. (Ed.). Current and selected bibliographies on benthic biology - 2014. Society for Freshwater Science, Logan, Utah. 237 pp.
• Jacobus, L. M. 2015. Pp. 35-41, in: Wetzel, M. J. (Ed.). Current and selected bibliographies on benthic biology Ñ 2013. Society for Freshwater Science, Logan, Utah. 301 pp.
• Kluge, N., Selvakumar, C., Sivaramakrishnan, K.G. & Jacobus, L.M. 2015. Contribution to the knowledge of the mayfly genus Teloganella Ulmer, 1939 (Ephemeroptera: Ephemerelloidea). Zootaxa 4028(2): 287-295.
• Jacobus, L. M. 2015. Book review: The Mayflies of Europe. American Entomologist 61(3): 190-191. Doi:10.1093/ae/tmv056.
• Poteat, M.D., Jacobus, L.M. & Buchwalter, D.B. 2015. The importance of retaining a phylogenetic perspective in traits-based community analyses. Freshwater Biology 60(7): 1330-1339, doi: 10.1111/fwb.12571.
• Jacobus, L.M. 2015. First report of the snow crane fly Chionea scita Walker, 1848 (Diptera: Tipuloidea: Limoniidae) from Indiana. Proceedings of the Indiana Academy of Science 123(1)(2014): 65-66.
• Jacobus, L.M. 2015. Ephemeroptera. Pp. 37-44, in: Wetzel, M. J. (Ed.). Current and selected bibliographies on benthic biology--2013 (for 2012). Society for Freshwater Science, Waco, Texas. 294 pp.
• Jacobus, L.M. 2015. 2013 Ephemeroptera Bibliography. The Mayfly Newsletter 18(2): 5-8.
• Jacobus, L.M., Salles, F.F., & Sartori, M. 2015. Zootaxa Ephemeroptera Editors' Annual Summary and Acknowledgments (2014). The Mayfly Newsletter 18(2): 3.
• Jacobus, L.M. 2015. Expansion and growth of Mayfly Central website. The Mayfly Newsletter 18(2): 2.
• Kluge, N. J., and L. M. Jacobus. 2015. Monochoroterpes, a replacement name for Monophyllus Kluge, 2012 (Insecta: Ephemeroptera), nec Monophyllus Leach, 1821 (Mammalia: Chiroptera). Zootaxa 3946(1): 150.
• Gorski, M.R., Fox, A.D.E., McQueen, J.I., and L.M. Jacobus. 2015. Plauditus cestus (Provonsha & McCafferty, 1982) (Insecta: Ephemeroptera: Baetidae): New records from Virginia and the Northwest Territories, with notes on color variation. Check List 11(1)1498: 1-3.
• Morse, J.C., McCafferty, W.P., Stark, B.P., Jacobus, L.M. (Eds.). 2015. Larvae of the Southeastern USA Mayfly, Stonefly and Caddisfly Species. Highlands Biological Foundation, Highlands, North Carolina. 434 pp. Unpublished workshop manual.
• Selvakumar, C., K. G. Sivaramakrishnan, Luke M. Jacobus, S. Janarthanan and M. Arumugam. 2014. Two new genera and five new species of Teloganodidae (Ephemeroptera) from South India. Zootaxa 3846(1): 87-104.
• Jacobus, L.M. and N. A. Wiersema. 2014. The genera Anafroptilum Kluge, 2011 and Neocloeon Traver, 1932, reinstated status, in North America, with remarks about the global composition of Centroptilum Eaton, 1869 (Ephemeroptera: Baetidae). Zootaxa 3814(3): 385-391.
• Lima, L. R. C., Luke M. Jacobus and U. Pinheiro. 2014. First description of imago and redescription of nymph for Cloeodes irvingi Waltz & McCafferty, 1987 (Ephemeroptera: Baetidae). Zootaxa 3838(5): 575-582.
• Jacobus, L. M. and F. F. Salles. 2014. Zootaxa Ephemeroptera Editors' Annual Summary and Acknowledgments (2013). The Mayfly Newsletter 18(1): 3.
• Jacobus, L.M. 2014. 2012 Ephemeroptera Bibliography. The Mayfly Newsletter 18(1): 6-10.
• Jacobus, L.M. 2014. New Facebook Page on Mayflies. The Mayfly Newsletter 18(1): 4.
• Jacobus, L. M., N. A. Wiersema and J. M. Webb. 2014. Identification of Far Northern and Western North American mayfly larvae (Insecta: Ephemeroptera), north of Mexico; Version 2. Joint Aquatic Science meeting, Portland, Oregon. 176 pp. + suppl. Unpublished workshop manual.Jacobus, L. M. 2013. 2011 Ephemeroptera bibliography. The Mayfly Newsletter 17(2): 8-10.
• Jacobus, L. M. 2013. A new junior synonym for the Holarctic species Ametropus fragilis Albarda, 1878 (Insecta: Ephemeroptera: Ametropodidae). Proceedings of the Indiana Academy of Science 122(1): 18-19.
• Boldrini, R., L. M. Jacobus, F. F. Salles and A. M. O. Pes. 2013. The adults of Camelobaetidius janae Dominique & Thomas, 2001, and C. yacutinga Nieto, 2003, with a new synonym for C. leentvaari Demoulin, 1966 (Ephemeroptera: Baetidae). Zootaxa 3702(2): 150-158.
• Jacobus, L. M. and J. M. Webb. 2013. A new junior synonym for Raptoheptagenia cruentata (Walsh, 1863) and remarks about Nearctic Heptagenia Walsh, 1863 (Insecta: Ephemeroptera: Heptageniidae). Proceedings of the Indiana Academy of Science 121(2): 143-146.
• Jacobus, L. M. 2013. South Carolina mayflies (Insecta: Ephemeroptera) of conservation concern. Journal of the South Carolina Academy of Science 11(1): 23-26. Available online: https://scholarcommons.sc.edu/jscas/vol11/iss1/5/
• Jacobus, L. M., N. A. Wiersema and J. M. Webb. 2013. Identification of Far Northern and Western North American mayfly larvae (Insecta: Ephemeroptera), north of Mexico. Southwest Association of Freshwater Invertebrate Taxonomists, California State University, Long Beach. 107 pp. + suppl. Unpublished workshop manual.
• Jacobus, L. M. 2013. A family key for the extant mayfly larvae of the world (Insecta: Ephemeroptera). Southwest Association of Freshwater Invertebrate Taxonomists, California State University, Long Beach. 29 pp. Unpublished workshop manual.
• Jacobus, L. M. 2013. Key to Ephemerellidae larvae of the Southeastern USA. Carolina Area Benthologists Association, Hot Springs, North Carolina. 25 pp. Unpublished workshop manual.
• Garlick, A., B. C. Kondratieff and L. M. Jacobus. 2012. Geographic range extension to Wyoming, USA, for Paraleptophlebia praepedita (Eaton, 1884) (Insecta, Ephemeroptera, Leptophlebiidae). Check List 8: 829-831. Available online: https://www.checklist.org.br/archive?vol=8&num=4
• Jacobus, L. M. 2012. South Carolina mayflies (Insecta: Ephemeroptera) of conservation concern. Report, Comprehensive Wildlife Conservation Strategy, South Carolina Department of Natural Resources, Columbia, South Carolina. 12 pp.
• Webb, J. M., L. M. Jacobus, D. H. Funk, X. Zhou, B. Kondratieff, C. J. Geraci, R. E. DeWalt, D. Baird, B. Richard, I. Phillips and P. D. N. Hebert. 2012. A DNA Barcode Library for North American Ephemeroptera: Progress and Prospects. PLoS ONE 7(5): e38063. doi: 10.1371/journal.pone.0038063, available online: https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0038063
• McCafferty, W. P., R. P. Randolph and L. M. Jacobus. 2012. Mayflies of the Intermountain West. Memoirs of the American Entomological Institute 85: i-iii + 1-317.
• Braasch, D., and L. M. Jacobus. 2011. Two new species of Afronurus Lestage, 1924, from Hong Kong, China (Ephemeroptera: Heptageniidae). Zootaxa 3062: 64-68, available online: https://www.mapress.com/zootaxa/2011/f/zt03062p068.pdf
• Shokralla, S., X. Zhou, D. H. Janzen, W. Hallwachs, J.-F. Landry, L. M. Jacobus and M. Hajibabaei. 2011. Pyrosequencing for mini-barcoding of fresh and old museum specimens. PLoS ONE 6(7): e21252. doi:10.1371/journal.pone.0021252, available online: https://www.plosone.org/article/info%3Adoi%2F10.1371%2Fjournal.pone.0021252
• Zhou, X., J. L. Robinson, C. J. Geraci, C. R. Parker, O. S. Flint, Jr., D. Etnier, D. Ruiter, R. E. DeWalt, L. M. Jacobus and P. D. N. Hebert. 2011. Accelerated construction of a regional DNA barcode reference library: Caddisflies (Trichoptera) in the Great Smoky Mountains National Park. Journal of the North American Benthological Society 30: 131-162 + 3 suppl.
• Jacobus, L. M. 2011. The mayfly family Baetidae in the Southeastern USA (Insecta: Ephemeroptera). Carolina Area Biologists Workshop, Hot Springs, North Carolina. 25 pp. Unpublished workshop manual.
• Jacobus, L. M. 2011. The mayfly family Baetidae in the Southeastern USA (Insecta: Ephemeroptera), revised edition. Unpublished workshop manual.
• Cook, K. R., A. L. Parks, L. M. Jacobus, T. C. Kaufman and K. A. Matthews. 2010. New research resources at the Bloomington Drosophila Stock Center. Fly 4: 87-90, available online: https://www.tandfonline.com/doi/abs/10.4161/fly.4.1.11230
• Hill, M. A., J. Pfeiffer and L. M. Jacobus. 2010. A new genus and new species of Baetidae (Ephemeroptera) from lakes and reservoirs in eastern North America. Zootaxa 2481: 61-68, available online: https://www.mapress.com/zootaxa/2010/f/zt02481p068.pdf
• Gattolliat, J.-L. and L. M. Jacobus. 2010. Madaechinopus, a replacement name for Echinopus Gattolliat, 2002 (Ephemeroptera: Baetidae), nec Fisher, 1814 (Monotremata: Tachyglossidae) and nec Schoenherr, 1836 (Coleoptera: Curculionidae). Aquatic Insects 32: 159-160.
• Zhou, X., L. M. Jacobus, R. E. DeWalt, S. J. Adamowicz and P. D. N. Hebert. 2010. The Ephemeroptera, Plecoptera, and Trichoptera fauna of Churchill (Manitoba, Canada): insights into biodiversity patterns from DNA barcoding. Journal of the North American Benthological Society 29: 814-837.
• Jacobus, L. M. and E. D. Fleek. 2010. Insecta, Ephemeroptera, Ephemerellidae, Attenella margarita (Needham, 1927): Southeastern range extension to North Carolina, USA. Check List 6: 311-313, available online: https://www.checklist.org.br/archive?vol=6&num=2
• Jacobus, L. M. 2010. Taxonomic review of the Caudatella heterocaudata (McDunnough) and C. hystrix (Traver) complexes (Insecta: Ephemeroptera: Ephemerellidae). Psyche 2010 (493935), available online: https://www.hindawi.com/journals/psyche/2010/493935.html
• McCafferty, W. P., D. R. Lenat, L. M. Jacobus and M. D. Meyer. 2010. The mayflies (Ephemeroptera) of the southeastern United States. Transactions of the American Entomological Society 136: 221-233.
• Ogden, T.H., J. T. Osborne, L. M. Jacobus and M. F. Whiting. 2009. Combined molecular and morphological phylogeny of Ephemerellinae (Ephemerellidae: Ephemeroptera), with remarks about classification. Zootaxa 1991: 28-42.
• Jacobus, L. M. and W. P. McCafferty. 2009. New synonym of Procloeon texanum McCafferty and Provonsha (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington 111: 286-287.
• Jacobus, L. M. and W. P. McCafferty. 2009. Two objective synonyms of Afrotropical mayfly genera (Ephemeroptera: Baetidae, Caenidae). Proceedings of the Entomological Society of Washington 111: 757.
• Jacobus, L. M. 2009. Insecta, Ephemeroptera, Ephemerellidae, Teloganopsis subsolana: Distribution extension and first report since its original description. Check List 5: 537-538, available online: https://www.checklist.org.br/archive?vol=5&num=3
• Jacobus, L. M., C.-F. Zhou and W. P. McCafferty. 2009. Two new species of Asian Serratella Edmunds (Ephemeroptera: Ephemerellidae). Zootaxa 2268: 52-58.
• Xie, H., Y.-Y. Jia, P. Chen, L. M. Jacobus and C.-F. Zhou. 2009. Two New Cincticostella species from China with a larval key to species of the genus (Ephemeroptera: Ephemerellidae). Zootaxa 2299: 53-61.
• Zhou, X., S. J. Adamowicz, L. M. Jacobus, R. E. DeWalt and P. D. N. Hebert. 2009. Towards a comprehensive barcode library for Arctic life: Ephemeroptera, Plecoptera and Trichoptera of Churchill, Manitoba, Canada. Frontiers in Zoology 6(30), available online: www.frontiersinzoology.com/content/6/1/30
• McCafferty, W. P. and L. M. Jacobus. 2008. Insecta, Ephemeroptera, Baetidae: Range extensions and new state records from Kansas, USA. Check List 4: 92-97, available online: https://www.checklist.org.br/archive?vol=4&num=2
• Jacobus, L. M. and W. P. McCafferty. 2008. Revision of Ephemerellidae Genera (Ephemeroptera). Transactions of the American Entomological Society 134: 185-274.
• Myers, L. W., L. M. Jacobus and B. C. Kondratieff. 2008. Insecta, Ephemeroptera: New and additional records from New York (U.S.A.). Check List 4: 415-423, available online: https://www.checklist.org.br/archive?vol=4&num=4
• McCafferty, W. P., L. M. Jacobus, J. M. Webb and M. D. Meyer. 2008. Insecta, Ephemeroptera: Range extensions and new records for Ontario and Canada. Check List 4: 449-452, available online: https://www.checklist.org.br/archive?vol=4&num=4
• Zhou, X., L. M. Jacobus and P. D. N. Hebert. 2008. DNA barcoding North American mayflies. The Mayfly Newsletter 15(2): 1-3.
• DeWalt, R. E., L. M. Jacobus and W. P. McCafferty. 2007. Summer Ephemeroptera, Plecoptera and Trichoptera of Southwestern Drainages in Great Smoky Mountains National Park. Proceedings of the Entomological Society of Washington 109: 136-154.
• Jacobus, L. M. and W. P. McCafferty. 2007. Reinstatement of Serratella serratoides and status of Ephemerella molita (Ephemeroptera: Ephemerellidae). Proceedings of the Entomological Society of Washington 109: 730-732.
• Jacobus, L. M., W. P. McCafferty, and R. W. Sites. 2007. A new species and stage associations in the Genus Crinitella (Ephemeroptera: Ephemerellidae). Zootaxa 1611: 45-53.
• Parker, C. R., O. S. Flint, Jr., L. M. Jacobus, B. C. Kondratieff, W. P. McCafferty, and J. C. Morse. 2007. Ephemeroptera, Plecoptera, Megaloptera, and Trichoptera of Great Smoky Mountains National Park. Southeastern Naturalist, Special Issue 1: 159-174.
• Sun, L., A. Sabo, M. D. Meyer, R. P. Randolph, L. M. Jacobus, W. P. McCafferty, and V. R. Ferris. 2006. Tests of Current Hypotheses of Mayfly (Ephemeroptera) Phylogeny by Using Molecular (18s rDNA) Data. Annals of the Entomological Society of America 99: 241-252.
• Jacobus, L. M., W. P. McCafferty, and J.-L. Gattolliat. 2006. Taxonomy of Afrotropical Securiops, new genus, and Cloeodes Traver (Ephemeroptera: Baetidae). African Entomology 14: 129-140.
• Jacobus, L. M., R. W. Newell, and W. P. McCafferty. 2006. First adult and egg descriptions of Caudatella edmundsi (Ephemeroptera: Ephemerellidae), from Montana (U.S.A.), with habitat notes. Entomological News 117: 175-180.
• Zhou, C.-F., L. M. Jacobus, and W. P. McCafferty. 2006. New Synonyms of Serratella jinghongensis (Ephemeroptera: Ephemerellidae). Entomological News 117: 237-238.
• Jacobus, L. M. and W. P. McCafferty. 2006. A new species of Acentrella Bengtsson (Ephemeroptera: Baetidae) from Great Smoky Mountains National Park, USA. Aquatic Insects 28: 101-111.
• Jacobus, L. M. and W. P. McCafferty. 2006. Phylogenetic revision of Ephemerythidae (Ephemeroptera: Pannota). Transactions of the American Entomological Society 132: 75-79.
• Jacobus, L. M. and W. P. McCafferty. 2006. Reevaluation of the phylogeny of the Ephemeroptera Infraorder Pannota (Furcatergalia), with adjustments to higher classification. Transactions of the American Entomological Society 132: 81-90, 429-430.
• Jacobus, L. M., and W. P. McCafferty. 2006. A New Oriental Species of Uracanthella (Ephemeroptera: Ephemerellidae) from Thailand and Vietnam. Entomological News117: 276-280.
• Jacobus, L. M., W. P. McCaffety. 2006. Notable Ephemeroptera Records (Baetidae, Leptophlebiidae, Metretopodidae) from Indiana. Entomological News 117: 344-346.
• McCafferty, W. P. and L. M. Jacobus. 2006. A new Oriental species of Behningia (Ephemeroptera: Behningiidae). Journal of Insect Science 6 (47), available online: https://academic.oup.com/jinsectscience/article/6/1/47/872085
• Jacobus, L. M. and W. P. McCafferty. 2005. Mayfly (Ephemeroptera) Diversity in the Great Smoky Mountains. Southeastern Biology 52(1): 45-46.
• Jacobus, L. M. and W. P. McCafferty. 2005. Validation of Potamocloeon macafertiorum Lugo-Ortiz (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington. 107: 474-475.
• Jacobus, L. M. and W. P. McCafferty. 2005. A new species and new synonym in Camelobaetidius (Ephemeroptera: Baetidae). Journal of the Kansas Entomological Society 78: 153-157.
• Randolph, R. P. and L. M. Jacobus. 2005. Northwest Biological Assessment Workgroup Taxonomic Workshop Manual: Ephemeroptera. Xerces Society, Portland, Oregon. 178 pp. Unpublished workshop manual.
• Jacobus, L. M., W. P. McCafferty, and R. W. Sites. 2005. A new synonym and new Thailand records of Cincticostella femorata (Tshernova) (Ephemeroptera: Ephemerellidae). Proceedings of the Entomological Society of Washington 107: 733-734.
• Jacobus, L. M. and W. P. McCafferty. 2005. Apobaetis futilis, a new combination in Nearctic Baetidae (Ephemeroptera). Proceedings of the Entomological Society of Washington 107: 979.
• McCafferty W. P.,  R. D. Waltz, J. M. Webb, and L. M. Jacobus. 2005. Revision of Heterocloeon McDunnough (Ephemeroptera: Baetidae). Journal of Insect Science 5 (35), available online: https://doi.org/10.1673/1536-2442(2005)5[1:ROHMEB]2.0.CO;2
• Jacobus, L. M., W. P. McCafferty, and R. W. Sites. 2005. Significant range extensions for Kangella and Vietnamella (Ephemeroptera: Ephemerellidae, Vietnamellidae). Entomological News 116: 268-270.
• Gattolliat, J.-L. and L. M. Jacobus. 2005. Review of the Dabulamanzia and Nesydemius (Ephemeroptera: Baetidae) species of Madagascar. Journal of the New York Entomological Society 113: 212-217.
•  Jacobus, L. M. and W. P. McCafferty. 2004. Contribution to the morphology and descriptive biology of Caurinella idahoensis (Ephemeroptera: Ephemerellidae). Western North American Naturalist 64: 101-108, available online: https://ojs.lib.byu.edu/wnan/index.php/wnan/article/view/1241/1098
• Jacobus, L. M. and W. P. McCafferty. 2004. First larval description of Paraleptophlebia calcarica Rowbotham and Allen (Ephemeroptera: Leptophlebiidae). Journal of the Kansas Entomological Society 77: 110-115.
• Jacobus, L. M., B. C. Kondratieff, M. D. Meyer, and W. P. McCafferty. 2004. Contribution to the biology and systematics of Ephemerella alleni (Ephemeroptera: Ephemerellidae). The Pan-Pacific Entomologist 79: 207-211.
• Jacobus, L. M. and M. Sartori. 2004. Review of the genus Hyrtanella (Ephemeroptera: Ephemerellidae). Zootaxa 785: 1-12, available online: https://www.mapress.com/zootaxa/2004f/zt00785.pdf
• McCafferty, W. P., M. D. Meyer, J. M. Webb, and L. M. Jacobus. 2004. New state and provincial records for North American small minnow mayflies (Ephemeroptera: Baetidae). Entomological News 115: 93-100.
• Jacobus, L. M. and W. P. McCafferty. 2004. Revisionary contributions to the genus Drunella (Ephemeroptera: Ephemerellidae). Journal of the New York Entomological Society 112: 127-147.
• Kluge, N. J., C. -F. Zhou, Luke M. Jacobus, W. P. McCafferty. 2004 [2005]. A new Southeast Asian mayfly species (Ephemeroptera: Ephemerellidae: Ephemerellinae). Journal of the New York Entomological Society 112: 148-152.
• Jacobus, L. M., C.-F. Zhou, W. P. McCafferty. 2004. Revisionary contributions to the genus Torleya (Ephemeroptera: Ephemerellidae). Journal of the New York Entomological Society 112: 153-175.
•  Jacobus, L. M. and W. P. McCafferty. 2003. A new synonym of Caudatella hystrix (Ephemeroptera: Ephemerellidae). Proceedings of the Entomological Society of Washington 105: 776-777.
• McCafferty, W. P., L. M. Jacobus, and T. Q. Wang. 2003. Reconfirmation of the genus Dentatella Allen (Ephemeroptera: Ephemerellidae). Proceedings of the Entomological Society of Washington 105: 786-788.
• Jacobus, L. M. and W. P. McCafferty. 2003. Revisions to the genus Crinitella (Ephemeroptera: Ephmerellidae). Journal of the New York Entomological Society 111: 48-50.
• Jacobus, L. M. and W. P. McCafferty. 2003. An introduction to the Smoky Mountain mayflies (Insecta: Ephemeroptera). Southeastern Biology 50: 358-360.
• Jacobus, L. M. and W. P. McCafferty. 2003. Revisionary contributions to North American Ephemerella and Serratella (Ephemeroptera: Ephemerellidae). Journal of the New York Entomological Society 111: 174-193.
• Jacobus, L. M. and W. P. McCafferty. 2002. Reinstatement of Rhithrogena manifesta Eaton (Ephemeroptera: Heptageniidae). Proceedings of the Entomological Society of Washington 104: 240-241.
• Jacobus, L. M. and W. P. McCafferty. 2002. Analysis of some historically unfamiliar Canadian mayflies (Ephemeroptera). The Canadian Entomologist 134: 141-155.
• Jacobus, L. M. and W. P. McCafferty. 2002. New synonyms for three North American Ephemeroptera species. Journal of the Kansas Entomological Society 75: 59-60.
• Randolph, R. P., W. P. McCafferty, D. Zaranko, L. M. Jacobus and J. M. Webb. 2002. New Canadian records of Baetidae (Ephemeroptera) and adjustments to North American Cloeon. Entomological News 113: 306-308.
• Jacobus, L. M. and W. P. McCafferty. 2002. Provancher’s quebecensis mayfly species (Ephemeroptera: Leptophlebiidae, Siphlonuridae). Entomological News 113: 315-317.
• Jacobus, L. M., W. P. McCafferty, and S. Spichiger. 2002. First adult description for Ephemerella hispida (Ephemeroptera: Ephemerellidae). Entomological News 113: 342-343.
• Jacobus, L. M. and W. P. McCafferty. 2001. New Ephemeroptera records from Pennsylvania. Entomological News 112: 144.
• Jacobus, L. M. and W. P. McCafferty. 2001. The mayfly fauna of New York State (Insecta: Ephemeroptera). Journal of the New York Entomological Society 109: 47-80.
• Jacobus, L. M. and W. P. McCafferty. 2001. Contribution to the systematics of Afrobaetodes Demoulin (Ephemeroptera: Baetidae). African Entomology 9: 97-103.
• Jacobus, L. M. and W. P. McCafferty. 2001. Adult description and commentary for two species of southeastern Nearctic Ephemerella (Ephemeroptera: Ephemerellidae). Entomological News 112: 294-298.
• McCafferty, W. P. and L. M. Jacobus. 2001. Revisions to Plauditus cestus and Plauditus gloveri (Ephemeroptera: Baetidae). Entomological News 112: 305-310.
• Jacobus, L. M. and W. P. McCafferty. 2001. Additions to the Canadian Ephemeroptera. Journal of the New York Entomological Society 109: 367-371.
• Jacobus, L. M. and W. P. McCafferty. 2000. Variability in the larvae of Serratella serrata (Ephemeroptera: Ephemerellidae). Entomological News 111: 39-44.